Both live on your account page (My Account in the header).
Sign-in email: choose Change email, enter the new address, and confirm it with the one-time code we send there. Your account switches over once the new address is verified.
Return address: this lives in your address book, not on the account page — it's an address, so that's where it sits. Under Your return addresses you can edit or remove any of the three you're allowed to save. We verify an address can receive mail when you save it. Remember it must be your own genuine, current address.
Your account page is where you choose which emails you get, under Communications:
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Mail handoff — email me when mail reaches the Postal Service.
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Mail delivery — email me when Certified Mail is delivered.
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Product updates — occasional email about new features. No spam.
All three are on to begin with, and you can change them whenever you like. If a letter runs into trouble we email you either way — that one isn't optional.